To start playing, press space or the play button in the wave panel. you can pause the spawning by clicking the pause button or by pressing space again. you have to kill 10 monsters from a wave to unlock the next wave. kill monster by clicking on them. once you earn enough gold, select a tower in the towers panel and then buy towers by clicking on any place of the walls in the game area. in the towers upgrades panel you can buy upgrades for your towers and unlock new ones.
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